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DPhil, University of Oxford.

BA(Hons) Biochemistry, University of Oxford.

Awards, Prizes or Honours

FMedSci, Academy of Medical Sciences, 2016.

EMBO Member, EMBO, 2017.

Editorial Boards

Journal of Cell Science, 1993.

Biochemical Journal, 1993.

External Committees

Executive Committee, Member, European Association for Cancer Research, 2010-2016.

Board of Directors, Director, Company of Biologists, 2011--.

British Society for Cell Biology, President, British Society for Cell Biology, 2006-2011.

Scientific Advisory Board, Chair, Wellcome Trust Centre for Cell:Matrix Research, University of Manchester, 2011-2016.

Molecular and Cellular Medicine Board (MCMB), Member, Medical Research Council, 2010-2014.

Career Catalyst Research Grants Committee, Member, Susan J Komen for the Cure, 2010-2010.

Senior and Career Development Fellowship Committee, Member, Cancer Research UK, 2006-2010.

Senior and Career Development Fellowship Committee, Chair, Medical Research Council (MRC), 2001-2006.

Molecular and Cellular Medicine Board (MCMB), Deputy Board Chair, Medical Research Council, 2012-2014.

Grants Committee, Member, Academy of Finland, 2012-2012.

Expert Review Group - Cell and Developmental Biology, Member, Wellcome Trust, 2012-2012.

Executive Committee, Secretary General, European Association of Cancer Research (EACR), 2014-2016.

Annual Meeting Scientific Program Committee, Committee Member, American Association of Cancer Research (AACR), 2014-2014.

Cancer Research Panel, Chair, Academy of Finland, 2014-2014.

New Investigator Awards Panel, Chair, Cancer Research UK, 2014--.

Science Committee, Member, Cancer Research UK, 2014-2020.

Scientific Advisory Board, Member, Wallenberg Centre for Molecular Translational Medicine, Gothenburg, Sweden, 2015-2020.

Scientific Advisory Board, Member, CRUK Manchester Institute, 2015-2017.

Scientific Advisory Board, Member, Join AZ-CRUK Functional Genomics Centre, 2019--.

Research Discovery Committee, Member, Cancer Research UK, 2020--.

Research Careers Committee, Chair, Cancer Research UK, 2020--.

International Advisory Board, Member, Oncode Institute, 2020--.